It was Williams' first appearance on Centre Court since the 2008 final, when she beat sister Serena for her second Wimbledon title in a row.
"I really enjoyed being out there," Venus said. "It's a special moment when you walk back as defending champion on that court."
Venus slipped five points into the match, one of several wobbly moments as she began her bid for a three-peat. She double-faulted in the opening game and had to erase two break points. She was passed the first two times she reached the net. She slipped and nearly fell a second time.
"It's grass," she said. "You're going to slip sometimes."
Williams found her footing, winning 14 consecutive points to help take a 5-1 lead. She had another spurt in the second set after losing serve for 2-all, and swept the final four games.
"Having won this title multiple times, you get that sense of what it takes to win," she said. "And I definitely have a good grip on that — what it takes to win this title."
Other players also took a tumble — Andy Roddick went down once during his victory, and even a ball boy fell on his face on Centre Court — and the biggest stumble was by Marat Safin. The two-time Grand Slam champion, who plans to retire at the end of the year, lost to qualifier Jesse Levine of the United States, 6-2, 3-6, 7-6 (4), 6-4.
"Not the way to finish Wimbledon story," Safin said. "But anyway, it's OK. That's life."
Levine, ranked 133rd, earned his first tour-level victory this year and his first win ever over a top-50 player.
"Marat's a great player, and I've watched him play growing up," said Levine, 21. "So it's a pretty surreal experience to be his last match at Wimbledon."
Attendance was 45,955, largest in tournament history.
Roddick followed Williams onto Centre Court and beat Jeremy Chardy 6-3, 7-6 (3), 4-6, 6-3. Roddick, seeded sixth, had only nine unforced errors and hit 47 winners, including 21 aces. He improved to 20-3 in tiebreakers this year.
Roddick was Wimbledon runner-up to Roger Federer in 2004 and 2005, but Andy Murray of Britain is considered the biggest obstacle for Federer this year.
"As far as who's talking about what, I don't really care," Roddick said. "I just want to go out and win matches."
The new retractable roof again worked well, keeping rain away for a second successive day. Play took place on a cloudless afternoon, prompting an official on the club's public-address system to urge that fans use sun block.
"It looks really nice, the roof," Williams said. "We haven't had to use it yet. It's kind of ironic. But I'm very sure it will get some use."
No. 13-seeded Ana Ivanovic, the 2008 French Open champion, saved two match points to beat Lucie Hradecka 5-7, 6-2, 8-6. This year's winner at Roland Garros, Svetlana Kuznetsova, defeated Akiko Morigami 6-3, 7-6 (1).
Top-ranked Dinara Safina opened another bid for her first Grand Slam title by beating Lourdes Dominguez Lino 7-5, 6-3. Safina said she was hampered by a sore left knee that has bothered her for two months, although she reached the French Open final less than three weeks ago.
Former No. 1 Jelena Jankovic beat Julia Goerges 6-4, 7-6 (0). Kimiko Date Krumm, a 38-year-old wild card who came out of retirement last year, lost in her first Wimbledon match since 1996 to No. 9-seeded Caroline Wozniacki, 5-7, 6-3, 6-1.
No. 17 Amelie Mauresmo, the 2006 champion, defeated Melinda Czink 6-1, 4-6, 6-2.
On the men's side, the third-seeded Murray began his bid to become the first British man to win Wimbledon since 1936 by beating American Robert Kendrick 7-5, 6-7 (3), 6-3, 6-4.
Juan Martin del Potro never faced a break point and swept Arnaud Clement 6-3, 6-1, 6-2. No. 12 Nikolay Davydenko beat Daniel Evans 6-2, 6-3, 6-3.
British wild card Alex Bogdanovic's record at Wimbledon fell to 0-8 when he lost to No. 20 Tomas Berdych 6-3, 6-4, 6-4.
Lleyton Hewitt, the 2002 champion, eased by American Robby Ginepri 6-4, 6-1, 6-1. Ginepri was bothered by a neck injury he suffered last week and received treatment from a trainer three times during the match.
The unseeded Hewitt and Federer are the only former champions in the men's draw. Hewitt plays del Potro next.
Williams prepared for Wimbledon as usual on hard courts back home in Florida, and didn't play a grass-court warmup tournament. But after her slow start she looked at home on the lawn.
In one game she smacked a backhand return up the line for a winner, then did the same thing from the other wing. Her second serve was unsteady, but she lost only six points on her first serve while hitting 29 winners and committing only 11 unforced errors.
"On the grass, I think you have the opportunity to make fantastic shots that are very entertaining and great plays," Williams said. "I think the game is more fast-paced. In a lot of ways, it makes it a lot more exciting."
Williams is only 6-4 since early April, but Wimbledon always brings her out of the doldrums. She's 51-4 at the All England Club since 2000, when she won the title for the first time. She's seeded third but the tournament favorite with London bookmakers.
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